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1 CRC Cell Cycle Genetics Group, University of Cambridge, Department of Genetics, Downing Site, Tennis Court Road, Cambridge CB2 3EH, UK
2 Department of Chemical Engineering, Instituto Superior Técnico, Av. Rovisco Pais, Lisboa 1049-001, and Cell Division Group, Instituto Gulbenkian de Ciencia, Rua Quinta Grande 6, Oeiras 2780-156, Portugal
3 School of Biological Sciences, 2.205 Stopford Building, University of Manchester, Oxford Road, Manchester M13 9PT, UK
4 Department of Cell Biology, Max-Planck-Institute of Biochemistry, Am Klopferspitz 18a, Martinsried D-82152, Germany
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